Changeset 479 for experiments/frams/foraminifera

Timestamp:

03/25/16 00:28:50 (9 years ago)

Author:

oriona

Message:

Parameters calibrated, diversification of initial population added, chamber growth and gametogenesis extended in time.

Location:

experiments/frams/foraminifera/data/scripts

Files:

• foraminifera.expdef (modified) (28 diffs)
• foraminifera.inc (modified) (5 diffs)

experiments/frams/foraminifera/data/scripts/foraminifera.expdef

@@ -23 +23 @@
 code:~
 
-
+global chambers;
+global colors;
+global dir_change;
+global max_chamber_volume;
+global movePerStep;
 global nutrientenergywaiting;
+global o;
 global reprocounter;
-global colors;
-global chambers;
-global o;
-global max_chamber_energ;
-global dir_change;
-global movePerStep;
 
 @include "foraminifera.inc"
@@ -83 +82 @@
 
         //time
-        ExpParams.secPerStep = 60;
+        ExpParams.secPerStep = 480;
         ExpParams.foramSpeedMmPerMin = 0.05;
         movePerStep = getMovePerStep();
@@ -92 +91 @@
 
         //reproduction
-        ExpParams.foramPop = 10;        
+        ExpParams.foramPop = 4; 
         ExpParams.crossprob = 0;
         ExpParams.mutationprob = 0;
-        ExpParams.repro_time = 20;
+        ExpParams.repro_time = 720;
+        ExpParams.gametoPeriod = 43200;
+        ExpParams.divisionCost = 15.6;
         reprocounter = 0;
+
+        init_chambers();
 
         //morphology
@@ -102 +105 @@
         ExpParams.zone1_range = micronsToFrams(1000);
         ExpParams.zone2_range = micronsToFrams(3000);
-        init_chambers();
+        ExpParams.chamber_growth_time = 720;
+        ExpParams.chamberCostPerSec = 0.000001;
         ExpParams.chamber_proculus_haplo = micronsToFrams(50);
         ExpParams.chamber_difference_haplo = 0.0;
         ExpParams.chamber_proculus_diplo = micronsToFrams(20);
         ExpParams.chamber_difference_diplo = 0.2;
-        max_chamber_energ = [Vector.new(), Vector.new()];
+
+        max_chamber_volume = [Vector.new(), Vector.new()];
         for (var j = 0; j < 2; j++)
         {
                 for (var i = 0; i < chambers[0].size; i++)
                 {
-                        max_chamber_energ[j].add(((energyFromVolume(getProperty(j, "chamber_proculus")) + energyFromVolume(getProperty(j, "chamber_proculus") + (i) * getProperty(j, "chamber_difference")))*(i+1))/2);
+                        max_chamber_volume[j].add(((volumeInMicrons(getProperty(j, "chamber_proculus")) + volumeInMicrons(getProperty(j, "chamber_proculus") + (i) * getProperty(j, "chamber_difference")))*(i+1))/2);  
                 }                                  
         }
@@ -120 +125 @@
         ExpParams.min_repro_energ_diplo = 6;
 
-        ExpParams.e_meta = 0.00005; 
-        ExpParams.energy_hib = 0.000025;
-        ExpParams.energy_move = 0.00005;
-
-        ExpParams.energies0_haplo = max_chamber_energ[0][0] - 0.001*max_chamber_energ[0][0];
-        ExpParams.energies0_diplo = max_chamber_energ[1][0] - 0.001*max_chamber_energ[1][0];
-        ExpParams.feedtrans = 0.125;
-        ExpParams.e_repro_cost_haplo = 0.7; 
-        ExpParams.e_repro_cost_diplo = 0.3;
+        ExpParams.e_meta = 0.0000005; 
+        ExpParams.energy_hib = 0.0000001;
+        ExpParams.energy_move = 0.0000005;
+
+        ExpParams.energies0_haplo = energyFromVolume(ExpParams.chamber_proculus_haplo,1);
+        ExpParams.energies0_diplo = energyFromVolume(micronsToFrams(1.25),1);
+        ExpParams.feedtrans = 0.001;
+        ExpParams.e_repro_cost_haplo = 0.3; 
+        ExpParams.e_repro_cost_diplo = 0.2;
+
+        ExpParams.e_death_level_haplo = 0.5;
+        ExpParams.e_death_level_diplo = 0.5;
 
         //nutrients
-        ExpParams.nutrientsize = micronsToFrams(10);
-        ExpParams.energy_nut = 100 * energyFromVolume(ExpParams.nutrientsize);
-        ExpParams.nutrientPop = 1;
-        ExpParams.feedrate = 100;
+        ExpParams.foodperiod = 480;
+        ExpParams.nutrientradius = micronsToFrams(10);
+        ExpParams.energy_nut = energyFromVolume(ExpParams.nutrientradius,1);
+        ExpParams.feedrate = 1;
+        ExpParams.ingestion = 0.25;
         nutrientenergywaiting = 0;
         ExpState.totaltestedcr = 0;
@@ -142 +151 @@
 @include "standard_placement.inc"
 
-function energyFromVolume(radius)
-{
-        return 4.0/3.0*Math.pi*Math.pow(radius,3);
+function volumeInMicrons(radiusInFrams)
+{
+        return 4.0/3.0*Math.pi*Math.pow(framsToMicrons(radiusInFrams),3);
+}
+
+function energyFromVolume(base, isRadiusInFrams)
+{
+        if (isRadiusInFrams == 1) //radius in frams
+        {
+                return ExpParams.picoCarbonPerMikro*volumeInMicrons(base);
+        }
+        else //volume in microns
+        {
+                return ExpParams.picoCarbonPerMikro * base;
+        }
 }
 
@@ -169 +190 @@
 }
 
-function addInitialForam(species, i)
-{
-        var geno = createForamGenotype(0, species, 0);
+function addForam(species, iter, chambernum, ploid)
+{
+        var geno = createForamGenotype(ploid, species, chambernum);
         var cr = Populations[0].add(geno);
-        cr.name = "Initial creature" + species + "_" + i;
+        cr.name = "Initial creature" + species + "_" + iter;
         placeCreatureRandomly(cr, 0, 0);
-        cr.energy0 = getProperty(0, "energies0");
+        cr.energy0 = energyFromVolume(max_chamber_volume[ploid][chambernum],0);
         cr.energy = cr.energy0;
         setGenotype({"opt" : 0, "cr" : cr, "species" : species});
+        if (ploid == 1)
+        {
+                cr.data->lifeparams->gen = 1;
+                cr.data->genes = [cr.data->genes, cr.data->genes]; //TODO two different genes sets
+        }
+}
+
+function addInitialForam(species, iter)
+{
+        var ploid = 0;
+        if (Math.rnd01 > 0.5)
+        {
+                ploid = 1;
+        }       
+        //add new foram with random energy bewtween starting energy and reproduction treshold
+        addForam(species, iter, Math.rndUni(0,getProperty(ploid, "min_repro_energ")),ploid);
 }
 
@@ -261 +298 @@
 // -------------------------------- foram begin -----------------------------------
 
-function setForamMeta(cr, gen)
-{
-        //cr.idleen = (ExpParams.e_meta * max_chamber_energ[gen][Math.min(lastChamberNum(cr), max_chamber_energ[gen].size-1)])*ExpParams.secPerStep;
+function setForamMeta(cr)
+{
+        //percent of current energy
         cr.idleen = (ExpParams.e_meta * cr.energy)*ExpParams.secPerStep;
 }
@@ -272 +309 @@
 }
 
+function getZoneRange(cr, zone_num)
+{
+        return ExpParams.["zone"+zone_num+"_range"];
+}
+
 function onForamsBorn(cr)
 {
-        setForamMeta(cr, 1);
+        setForamMeta(cr);
         if (ExpParams.visualize == 1)
         {
-                var ret = Populations[2].add("//0\np:sh=3,sx=0.01,sy="+ExpParams.zone1_range+",sz="+ExpParams.zone1_range+",ry=1.57,vr=1.0,1.0,1.0");
+                var ret = Populations[2].add("//0\np:sh=3,sx=0.01,sy="+getZoneRange(cr,1)+",sz="+getZoneRange(cr,1)+",ry=1.57,vr=1.0,1.0,1.0");
                 cr.data->reticulopodiacreature = ret;
         }
@@ -322 +364 @@
                 setGenotype({"cr" : cr2, "parent_genes" : cr.data->genes, "parent_lifeparams" : cr.data->lifeparams, "opt" : 2});
                 cr2.moveAbs(cr.center_x - cr2.size_x / 2, cr.center_y - cr2.size_y / 2, cr.pos_z);
-                setForamMeta(cr2, cr2.data->lifeparams->gen);
+                setForamMeta(cr2);
 
                 if (visualization(cr))
@@ -335 +377 @@
 {
         var p = cr.getMechPart(0);
-        var n = cr.signals.receiveSet("nutrient", ExpParams.zone2_range);
+        var n = cr.signals.receiveSet("nutrient", getZoneRange(cr,2));
 
         //if signals are received find the source of the nearest
@@ -354 +396 @@
                         distvec.sub(p.pos);
                         dist = distvec.length;
-                        if (dist < ExpParams.zone1_range)
+                        if (dist < getZoneRange(cr,1))
                         {
                                 if (n[i].value != null)
@@ -390 +432 @@
         if (cr.data->lifeparams->hibernated == 0)
         {
-                //cr.energy_m += (ExpParams.energy_move * max_chamber_energ[cr.data->lifeparams->gen][Math.min(lastChamberNum(cr), (max_chamber_energ[cr.data->lifeparams->gen].size)-1)])*ExpParams.secPerStep;
-                cr.energy_m += (ExpParams.energy_move * cr.energy)*ExpParams.secPerStep;
+                //percent of maximal energy
+                cr.energy_m += (ExpParams.energy_move * cr.data->lifeparams->max_energy_level)*ExpParams.secPerStep;
         }
 }
@@ -422 +464 @@
         }
         //random move
-        else if (cr.lifespan%(dir_change/ExpParams.secPerStep) == 0)
+        else if (cr.lifespan%int(dir_change/ExpParams.secPerStep) == 0)
         {
                 cr.data->lifeparams->dir = randomDir();
@@ -445 +487 @@
 {
         cr1.localDrive = XYZ.new(0,0,0);
-        var e = ExpParams.feedtrans*ExpParams.secPerStep; //TODO efficiency dependent on age
-        e = Math.min(cr2.energy, e) + 0.0000001;
-        //Simulator.print("transferring "+e+" to "+cr1.name+" from "+cr2.name+" ("+cr2.energy+")");
-        cr2.energy_m = cr2.energy_m + e;
-        cr1.energy_p = cr1.energy_p + e;
+        var e =  ExpParams.feedtrans*cr1.energy;//ExpParams.feedtrans*cr1.energy*ExpParams.ingestion*ExpParams.secPerStep; //TODO efficiency dependent on age
+        e = Math.min(cr2.energy, e);
+        e *= ExpParams.secPerStep;
+        //Simulator.print("transferring "+e +"("+e*ExpParams.ingestion+")"+" to "+cr1.name +" ("+ cr1.energy+") " +" from "+cr2.name+" ("+cr2.energy+") "+ e/ExpParams.secPerStep+ " per sec");
+        cr2.energy_m = cr2.energy_m + e + 0.0000001;
+        cr1.energy_p = cr1.energy_p + e*ExpParams.ingestion;
         if (cr1.data->lifeparams->hibernated == 1)
         {
@@ -460 +503 @@
         if (cr.data->lifeparams->hibernated == 1)
         {
-                setForamMeta(cr, cr.data->lifeparams->gen); //unhibernate
+                setForamMeta(cr); //unhibernate
         } 
         else
         {
-                //cr.idleen = (ExpParams.energy_hib * max_chamber_energ[cr.data->lifeparams->gen][Math.min(lastChamberNum(cr), (max_chamber_energ[cr.data->lifeparams->gen].size)-1)])*ExpParams.secPerStep; //hibernate
                 cr.idleen = (ExpParams.energy_hib * cr.energy)*ExpParams.secPerStep; //hibernate
         }
@@ -470 +512 @@
 }
 
+
 function onForamsStep(cr)
 {
-        cr.getMechPart(0).orient.set(o);
-        if (visualization(cr))
-        {
-                cr.data->reticulopodiacreature.moveAbs(cr.center_x-ExpParams.zone1_range, cr.center_y-ExpParams.zone1_range, cr.center_z-ExpParams.zone1_range-getProperty(cr.data->lifeparams->gen, "chamber_proculus"));
-        }
-
-        if (deathConditions(cr) == 1)
-        {
-                Populations[0].kill(cr);
-                return;
-        }
-
-        foramMove(cr);
-
-        var repro = foramReproduce(cr);
-        if (repro == 1)
-        {
-                return;
-        }
-
-        cr.data->lifeparams->max_energy_level = Math.max(cr.energy, cr.data->lifeparams->max_energy_level);
-        if  (lastChamberNum(cr) <= chambers[0].size-1)
-        {
-                if ((cr.data->lifeparams->max_energy_level >= max_chamber_energ[cr.data->lifeparams->gen][lastChamberNum(cr)]))         
-                {
-                        foramGrow(cr, lastChamberNum(cr)); 
-                }       
+        //checking for gametogenesis process
+        if (cr.data->lifeparams->division_time > 0)
+        {
+                cr.data->lifeparams->division_time = Math.max(cr.data->lifeparams->division_time-1,0);
+        }
+        //checking for end of gametogenesis
+        else if (cr.data->lifeparams->division_time == 0)
+        {
+                //waiting for gamets fusion
+        }
+        //checking for chamber growth process
+        else if (cr.data->lifeparams->chamber_growth > 0)
+        {
+                cr.data->lifeparams->chamber_growth = Math.max(cr.data->lifeparams->chamber_growth-1,0);
+                //Simulator.print("chamber growing, time left = " + cr.data->lifeparams->chamber_growth*ExpParams.secPerStep);
+                cr.energy_m += ExpParams.chamberCostPerSec * cr.energy * ExpParams.secPerStep;
+
+                //Simulator.print("energy " + cr2.energy + " subtracting " + growth_cost);
+        }
+        //checking for end of chamber growth process
+        else if (cr.data->lifeparams->chamber_growth == 0)
+        {
+                foramGrow(cr, lastChamberNum(cr)); 
+                cr.data->lifeparams->chamber_growth = -1;
+                //Simulator.print("chamber "+ (lastChamberNum(cr) + 1) +" complete");
+        }
+        else
+        {
+                //update of metabolism rate
+                if (cr.data->lifeparams->hibernated == 0)
+                {
+                        setForamMeta(cr);
+                }
+
+                cr.getMechPart(0).orient.set(o);
+                if (visualization(cr))
+                {
+                        cr.data->reticulopodiacreature.moveAbs(cr.center_x-getZoneRange(cr,1), cr.center_y-getZoneRange(cr,1), cr.center_z-getZoneRange(cr,1)-getProperty(cr.data->lifeparams->gen, "chamber_proculus"));
+                }
+
+                if (deathConditions(cr) == 1)
+                {
+                        Populations[0].kill(cr);
+                        return;
+                }
+
+                foramMove(cr);
+
+                var repro = foramReproduce(cr);
+                if (repro == 1)
+                {
+                        return;
+                }
+
+                cr.data->lifeparams->max_energy_level = Math.max(cr.energy, cr.data->lifeparams->max_energy_level);
+
+                //cheking conditions of chamber growth process start
+                if  (lastChamberNum(cr) != chambers[0].size-1)
+                {
+                        if ((cr.data->lifeparams->max_energy_level >= energyFromVolume(max_chamber_volume[cr.data->lifeparams->gen][lastChamberNum(cr)],0)))    
+                        {
+                                cr.data->lifeparams->chamber_growth = int(ExpParams.chamber_growth_time/ExpParams.secPerStep);
+                        }       
+                }
         }       
 }
@@ -504 +584 @@
 function deathConditions(cr)
 {
-        if ((cr.energy <= getProperty(cr.data->lifeparams->species,"e_death_level")) || (Math.rnd01 < ExpParams.hunted_prob))
+        if ((cr.energy <= getProperty(cr.data->lifeparams->gen,"e_death_level")*cr.data->lifeparams->max_energy_level) || (Math.rnd01 < ExpParams.hunted_prob))
+        {
                 return 1;
+        }
         else
                 return 0;
@@ -528 +610 @@
 // -------------------------------- nutrient begin --------------------------------
 
-function createNutrientGenotype(nutrientsize, zone1_range)
-{
-        return "//0\np:sh=3,sx="+nutrientsize+",sy="+nutrientsize+",sz="+nutrientsize+",ry=1.5,vr=0.0,1.0,0.0";
+function createNutrientGenotype(nutrientradius)
+{
+        return "//0\np:sh=3,sx="+nutrientradius+",sy="+nutrientradius+",sz="+nutrientradius+",ry=1.5,vr=0.0,1.0,0.0";
 }
 
@@ -540 +622 @@
 function addNutrient()
 {
-        var cr = Populations[1].add(createNutrientGenotype(ExpParams.nutrientsize, ExpParams.zone1_range));
+        var cr = Populations[1].add(createNutrientGenotype(ExpParams.nutrientradius));
 
         cr.name = "Nutrients";
@@ -553 +635 @@
         if (ExpParams.visualize == 1)
         {
-                var nutsize = ExpParams.nutrientsize*10;
+                var nutsize = ExpParams.nutrientradius*10;
                 var nut = Populations[2].add("//0\np:sh=2,sx="+nutsize+",sy="+nutsize+",sz="+nutsize+",ry=1.5,vr=0.0,1.0,0.0");
                 cr.data->reticulopodiacreature = nut;
@@ -571 +653 @@
 {
         nutrientenergywaiting = nutrientenergywaiting + 1;
-        if (nutrientenergywaiting > ExpParams.feedrate/ExpParams.secPerStep)
-        {
-                for (var i = 0; i < ExpParams.nutrientPop; i++)
+        if (nutrientenergywaiting*ExpParams.secPerStep >= ExpParams.foodperiod)
+        {
+                for (var i = 0; i < ExpParams.feedrate*(ExpParams.secPerStep/ExpParams.foodperiod); i++)
                 {    
                         addNutrient();
@@ -668 +750 @@
 
 // -------------------------------- step end --------------------------------
+//TODO default params values in frams instead of microns/seconds
 
 @include "standard_events.inc"
@@ -692 +775 @@
 
 prop:
+id:gametSuccessRate
+name:Ratio of successful gamets
+type:f 0.001
+group:Foraminifera
+
+prop:
+id:gametoPeriod
+name:Time of gametogenesis
+type:f 720
+group:Foraminifera
+
+prop:
+id:picoCarbonPerMikro
+name:Picograms of carbon in cubed micrometer
+type:f 0.13
+group:Foraminifera
+
+prop:
 id:secPerStep
 name:Seconds per simulation step
@@ -705 +806 @@
 
 prop:
+id:divisionCost
+name:Cost of division in pG
+type:f
+group:Foraminifera
+
+prop:
 id:e_repro_cost_diplo
 name:Cost of reproduction
 type:f 0.1 0.9 0.3
+group:Foraminifera
+
+prop:
+id:chamber_growth_time
+name:Time of the chamber growth in seconds
+type:f
+group:Foraminifera
+
+prop:
+id:chamberCostPerSec
+name:Cost of growning chamber per second
+type:f
 group:Foraminifera
 
@@ -791 +910 @@
 id:e_death_level_haplo
 name:Minimal level of energy to sustain life of haploid
-type:f 0 20 4
+type:f 0 20 0.2
 group:Foraminifera
 
@@ -797 +916 @@
 id:e_death_level_diplo
 name:Minimal level of energy to sustain life of diploid
-type:f 0 20 0.25
+type:f 0 20 0.2
 group:Foraminifera
 
@@ -857 +976 @@
 
 prop:
+id:ingestion
+name:Ingestion rate
+type:f
+group:Energy
+
+prop:
 id:energy_nut
 name:Nutrient energy
-type:f 0 1000
+type:f 0 100000
 group:Energy
 
 prop:
 id:feedtrans
-name:Ingestion multiplier
-type:f 0 100
+name:Energy transfer per second
+type:f 0 100000
 group:Energy
 
 prop:
-id:nutrientsize
+id:foodperiod
+name:Time between food occurrences
+type:f 0 100000
+group:Energy
+
+prop:
+id:nutrientradius
 name:Nutrient size
 type:f 0.1 0.9 0.1
 group:Energy
-
-prop:
-id:nutrientPop
-name:Nutrient population size
-group:Energy
-type:d 1 1000 10
 
 prop:

experiments/frams/foraminifera/data/scripts/foraminifera.inc

@@ -55 +55 @@
         if (mode->opt == 0) //initial
         {
-                mode->cr.data->genes = {"min_repro_energies" : [max_chamber_energ[0][getProperty(0, "min_repro_energ")], max_chamber_energ[1][getProperty(1, "min_repro_energ")]], "hibernation" : mode->species};
-                mode->cr.data->lifeparams = {"max_energy_level" : getProperty(0,"energies0"), "gen" : 0,  "hibernated" : 0, "species" : mode->species, "reproduce" : 0, "dir" : randomDir()};
+                mode->cr.data->genes = {"min_repro_energies" : [energyFromVolume(max_chamber_volume[0][getProperty(0, "min_repro_energ")],0), energyFromVolume(max_chamber_volume[1][getProperty(1, "min_repro_energ")],0)], "hibernation" : mode->species};
+                mode->cr.data->lifeparams = {"max_energy_level" : getProperty(0,"energies0"), "gen" : 0,  "hibernated" : 0, "species" : mode->species, "reproduce" : 0, "dir" : randomDir(), "chamber_growth" : -1, "division_time" : -1};
         }
         else if (mode->opt  == 1) //child
         {
-                mode->cr.data->lifeparams = {"max_energy_level" : getProperty(1 - mode->parent_lifeparams->gen,"energies0"), "gen" : 1 - mode->parent_lifeparams->gen,  "hibernated" : 0, "species" : mode->parent_lifeparams->species, "reproduce" : 0, "dir" : randomDir()};
+                mode->cr.data->lifeparams = {"max_energy_level" : getProperty(1 - mode->parent_lifeparams->gen,"energies0"), "gen" : 1 - mode->parent_lifeparams->gen,  "hibernated" : 0, "species" : mode->parent_lifeparams->species, "reproduce" : 0, "dir" : randomDir(), "chamber_growth" : -1, "division_time" : -1};
                 mode->cr.data->genes = mode->parent_genes;
         }
@@ -68 +68 @@
                 mode->cr.data->lifeparams = mode->parent_lifeparams;
         }
+}
+
+function gametsDivision(parent_energy, energy0)
+{
+        var number = 1;
+        var result = parent_energy;
+        while ((result-ExpParams.divisionCost) >= energy0)
+        {
+                result = (result-ExpParams.divisionCost)/2;
+                number *= 2; 
+        }
+        //Simulator.print("parent: " + parent_energy + " result: " + result + " number " + number);
+        return {"energy" : result, "number" : number};
 }
 
@@ -75 +88 @@
         if (clone == 1)
         {
-                energy0 = getProperty(0,"energies0");
-                number = (( 1 - getProperty(1, "e_repro_cost")) * parent.energy) / energy0;
+                var offspring = gametsDivision(parent.energy,getProperty(0,"energies0"));
+                energy0 = offspring->energy;
+                number = offspring->number;
                 new_genes = parent.data->genes;
                 parent.data->lifeparams->gen = 1 - parent.data->lifeparams->gen; //because of reversal of "gen" in createOffspring function
@@ -83 +97 @@
         else
         {
-                energy0 = getProperty(1,"energies0"); 
-                number = (((1 - getProperty(parent.data->lifeparams->gen, "e_repro_cost")) * parent.energy) + ((1 -(getProperty(parent.data->lifeparams->gen, "e_repro_cost"))) * parent2.energy)) / energy0;
+                var offspring1 = gametsDivision(parent.energy,getProperty(1,"energies0"));
+                var offspring2 = gametsDivision(parent2.energy,getProperty(1,"energies0"));
+                energy0 = (offspring1->energy+offspring2->energy); 
+                number = ExpParams.gametSuccessRate*(offspring1->number+offspring2->number)/2;
                 new_genes = [parent.data->genes, parent2.data->genes];
                 gen = 1 - parent.data->lifeparams->gen;
@@ -152 +168 @@
                                 if (parent1 != null && parent2 != null)
                                 {
-                                        reproduce_haploid(parent1, parent2, 0);
-                                        print_repro_info(parent1);
-                                        print_repro_info(parent2);
-                                        pop.kill(parent1);
-                                        pop.kill(parent2);
-                                        parent1 = null;
-                                        parent2 = null;
+                                        //when parents are ready for reproduction start gametogenesis
+                                        if (parent1.data->lifeparams->division_time == -1 && parent2.data->lifeparams->division_time == -1)
+                                        {
+                                                var time = int(ExpParams.gametoPeriod/ExpParams.secPerStep);
+                                                parent1.data->lifeparams->division_time = time;
+                                                parent2.data->lifeparams->division_time = time;
+                                                parent1.idleen = 0;
+                                                parent2.idleen = 0;
+                                                //Simulator.print("parents "+parent1.uid + " " + parent2.uid + " ready to repro: "+Simulator.stepNumber);
+                                        }
+                                        //when gametogenesis is finished fuse gamets 
+                                        else if (parent1.data->lifeparams->division_time == 0 && parent2.data->lifeparams->division_time == 0)
+                                        {
+                                                reproduce_haploid(parent1, parent2, 0);
+                                                print_repro_info(parent1);
+                                                print_repro_info(parent2);
+                                                //Simulator.print("parents "+parent1.uid + " " + parent2.uid + " reproduced: "+Simulator.stepNumber);
+                                                pop.kill(parent1);
+                                                pop.kill(parent2);
+                                                parent1 = null;
+                                                parent2 = null;
+                                                
+                                        }
                                 }       
                         }